| Ecology |
woodland (Kinki Research Group of Carabid Beetles 1979), tobacco fieleds (Kinki Research Group of Carabid Beetles 1979), cedar forests (Kinki Research Group of Carabid Beetles 1979, Togashi and Sugie 1994, Hiramatsu 2004), moist environment with little sunshine in low mountain range (East Japan Research Group of Carabid Beetles 1989), broad-leaved deciduous forest (Togashi and Hashimoto 1994), edge of broad-leaved deciduous forest neighboring paddy fields (Togashi and Hashimoto 1994), mountain (Kinki Research Group of Carabid Beetles 1979, Sota 1985c, Yahiro et al. 2001, Carabidological Society of Shiga 2003, Yahiro 2008), natural forests or near natural forests (Yahiro et al. 2001, Carabidological Society of Shiga 2003), dried secondary forest (Yahiro et al. 2001, Carabidological Society of Shiga 2003), beech forest (Kinki Research Group of Carabid Beetles 1979, Hiramatsu 2004), grassland (Hiramatsu 2004), adult (Sota 1985a, Sota 1985c), from spring to summer (Sota 1985a), from late May to late July (East Japan Research Group of Carabid Beetles 1989), from May to early August (Sota 1985c), larvae of Lepidoptera and Diptera (larva) (Sota 1985a, Sota 1987b), insect larva (larva) (Sota 1985c), hibernating in soil (East Japan Research Group of Carabid Beetles 1989), hibernating in decayed wood (East Japan Research Group of Carabid Beetles 1989) |
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