| Ecology |
mountain (Inoue 1953x), broad leaved deciduous forest (Higashi et al. 1983), margin of deciduous broad-leaved forest dominated (Higashi et al. 1983), coniferous forest (Higashi et al. 1983), margin of coniferous forest (Higashi et al. 1983), plantation of Larix leptolepis saplings (Higashi et al. 1983), plantation's margin of Larix leptolepis saplings (Higashi et al. 1983), Sasa nipponica bush left in a pasture (Higashi et al. 1983), from subalpine zone to alpine zone (East Japan Research Group of Carabid Beetles 1989), flatland in Hokkaido (East Japan Research Group of Carabid Beetles 1989), forest (East Japan Research Group of Carabid Beetles 1989, Kimoto and Yasuda 1995), Pinus pumila community (Kimoto and Yasuda 1991, Kimoto and Yasuda 1995), denuded snow-patch vegetation of the alpine region (Kimoto and Yasuda 1995), snowy wet grassland of the alpine region (Kimoto and Yasuda 1990, Kimoto and Yasuda 1992a, Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), Betula ermanii forest (Kimoto and Yasuda 1990, Kimoto and Yasuda 1991, Kimoto and Yasuda 1992a, 1992b, Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), Picea jezoensis and Betula ermanii forest (Kimoto and Yasuda 1991), higher mixed forest (Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), coniferous forest (Kimoto and Yasuda 1992a, 1992b, Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), lower mixed forest (Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), windy bare ground in the alpine region (Kimoto and Yasuda 1991, Kimoto and Yasuda 1995), broad-leaved forest in low mountain region (Kimoto and Yasuda 1991, Kimoto and Yasuda 1995), Alnus maximowiczii forest(Kimoto and Yasuda 1992a, Kimoto and Yasuda 1995), altherbosa (Kimoto and Yasuda 1995), Abies sachalinensis forest (Kimoto and Yasuda 1995, Hori 2003), Sasa kurilensis community (Kimoto and Yasuda 1995), Polygonum sachalinense community (Kimoto and Yasuda 1995), Quercus crispula forest (Kimoto and Yasuda 1995), mixed forest (Hori 2003), Picea abies forest (Hori 2003), broad-leaved secondary forest (Hori 2003), park forest (Hori 2003), adult (Inoue 1953x), early June (Inoue 1953x), carnivorous (Inoue 1953x), Lepidopteran larva (Inoue 1953x), a species of Melanoplinae (East Japan Research Group of Carabid Beetles 1989), hibernating within the decayed wood (Inoue 1953x) |
| 
