| Ecology |
mountainous area (Yamaguchi museum 1988), vineyard (Yano et al. 1989), forest adjoinig agroecosystems (Yahiro et al. 1990), paddy field (Yahiro et al. 1992), secondary forest of Alnus sieboldiana (Ishitani 1996), secondary forest of Pinus densiflora (Ishitani 1996), adult (Ishitani 1996), from April to June (Ishitani 1996), carnivorous (Ishitani 1996), Megascolecoidea (Kinki Research Group of Carabid Beetles 1979), hibernating in slope (Suga 1981), hibernating in decayed tree (Suga 1981), proovigenic type (Ishitani 1996), mainly hibernating in soil and rarely in decayed wood or under fallen tree (East Japan Research Group of Carabid Beetles 1989) |
| 
