Ecology |
riverbed (East Japan Research Group of Carabid Beetles 1989), light broad leaved forest (East Japan Research Group of Carabid Beetles 1989), from low land to subalpine zone (East Japan Research Group of Carabid Beetles 1989), secondary forest (East Japan Research Group of Carabid Beetles 1989), mountain (Hosoda 1996), coppice forest (Hosoda 1996, Hosoda et al. 1996), secondary forest (Hosoda 1996), primary forest (Hosoda 1996), broad-leaved forest in low mountain region (Kimoto and Yasuda 1991, Kimoto and Yasuda 1995), The larvae mainly eat Megascolecoidea (East Japan Research Group of Carabid Beetles 1989), mainly hibernating in soil and rarely in decayed wood or under fallen tree (East Japan Research Group of Carabid Beetles 1989) |
Remarks |
subspecies: ssp. albrechti, ssp. tohokuensis, ssp. esakianus, ssp. freyi, ssp. okumurai, ssp. hidakanus, ssp. itoi, ssp. echigo, ssp. awashimae, ssp. yamauchii ssp. haga, ssp. tusukubanus |