| Ecology |
forest, grassland and cultivated land etc. (Inoue 1953k), wilderness (East Japan Research Group of Carabid Beetles 1989), moor land (East Japan Research Group of Carabid Beetles 1989), forested land (East Japan Research Group of Carabid Beetles 1989), bareland (Higashi et al. 1983), grassland (Higashi et al. 1983), deciduous broad-leaved forest (Higashi et al. 1983), margin of deciduous broad-leaved forest dominated (Higashi et al. 1983), margin of coniferous forest (Higashi et al. 1983), plantation of L. leptolepis saplings (Higashi et al. 1983), plantation's margin of L. leptolepis saplings (Higashi et al. 1983), margin of alder thicket in wetlands (Higashi et al. 1983), high moor (Kimoto and Yasuda 1995), mountainous forest (Kimoto and Yasuda 1995), Betula ermanii forest (Kimoto and Yasuda 1992a, Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), lower mixed forest (Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), coniferous forest (Kimoto and Yasuda 1992a, 1992b, Kimoto and Yasuda 1995), Pinus pumila community (Kimoto and Yasuda 1995), Abies sachalinensis forest (Kimoto and Yasuda 1995), Sasa kurilensis community (Kimoto and Yasuda 1995), Polygonum sachalinense community (Kimoto and Yasuda 1995), riverain grassland (Kimoto and Yasuda 1995), peat bog (Kimoto and Yasuda 1995), Sasa palmata community (Kimoto and Yasuda 1995), high moor (Kimoto and Yasuda 1995), marshy forest (Kimoto and Yasuda 1995), Larix kaempferi forest (Hori 2003), woods (Hori 2003), adult (Inoue 1953k), June (Inoue 1953k), carnivorous (Inoue 1953k), once annually (Inoue 1953k), hibernating in soil (East Japan Research Group of Carabid Beetles 1989) |
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