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SNAME  Harpalus tridens Morawitz, 1862
JNAME  kogomokumushi
Collection ground Fukushima cho, Hokkaido Pref. 1995/8/5
Distribution Hokkaido, Honshu, Shikoku, Kyushu, Ryukyu
Ecology flatland (Ishitani 1996), paddy field (Torii 1974, Yahiro et al. 1992), grassland firm field in flatland (Inoue 1977e), riverbed (Terada 1983, Ishii et al. 1996, Ishitani et al. 1997, Yamazaki et al. 1999, Lee and Ishii 2009, 2010), crop field (Inoue and Hanada 1979), apple orchard (Koyama and Yamada 1980), bank (Togashi 1986, Aono 1987, Aono 1988), vineyard (Yano et al. 1989, Togashi and Ohbata 1995), forest adjoinig agroecosystems (Yahiro et al. 1990), forest edge (Sunose 1992, Sunose and Kurosawa 1992), slope of mound (Sunose 1992, Sunose and Kurosawa 1992), fallow field (Sunose 1992, Sunose and Kurosawa 1992), pear orchard (Togashi and Ura 1994), fig orchard (Ishitani and Yano 1994, Ishitani 1996), forage crop field (Ishitani et al. 1994), secondary forest of Pinus densiflora (Ishitani 1996), riverbed (Ishitani 1996), rape field (Ishitani 1996), field of residential area (Ishitani 1996), residential area (Ishitani 1996), broad-leaved secondary forest (Hori 2003), upland field (Hiramatsu 2004), vegetable field (Siddiquee and Nakamura 2004), field (Salah et al. 2004), broad leaved forest (Matsumoto 2005), grassland (Taniwaki et al. 2005), paddy field (Yahiro 2007, Kagawa 2008, Lee et al. 2008, Lee and Ishii 2009), coppice forest (Taniwaki et al. 2005, Kagawa 2008), forest (Suttiprapan and Nakamura 2007), urban park (Lee and Ishii 2009), Satoyama forest (Lee and Ishii 2009), adult and larva (Ishitani et al. 1994), larva (Ishitani 1996), adult (Yamazaki et al. 1999), from September to October (Ishitani 1996), autumn (Yamazaki et al. 1999), omnivorous (Ishitani 1996), this species is a natural enemy of Leucania separata Walker (Syojima 1905), it is injurious to the seeds of wheat and barley (Matsumura 1915, Shiraki 1954), the adults climb the stem of cultivated Polygonum ptatense Linne and feed on its seeds at night (Yago 1939), they injure the young root of graminaceous plant (Shiraki 1940), plant seeds (Inoue 1952e), collected by light trapping (Inoue and Hanada 1979, Aono 1988, Yahiro 1997, Suttiprapan and Nakamura 2007, Yahiro 2007), synovigenic type (Ishitani 1996)
Types 　 　 　
Remarks Harpalus tridens Morawitz, 1862, Mel. Biol., 4: 245 (Japan: “Hakodade”) (Habu 1973d), Harpalus tridens Morawitz, Ophonus (Pardileus) tridens Morawitz, Pseudophonus tridens Morawitz, Harpalus (Oponus) tridens Morawitz, Harpalus (Pardileus) Pecirkai Jedlicka, Pardileus tridens Morawitz, Ophonus tridens Morawitz, Harpalus (Pseudoophonus) tridens Morawitz (Habu 1973d)