| Ecology |
arboreal (East Japan Research Group of Carabid Beetles 1989, Suga 1981), flat woods or mountain (Inoue 1953d, East Japan Research Group of Carabid Beetles 1989), Quercus acutissima forest (Kinki Research Group of Carabid Beetles 1979), beech forest and forest margin (East Japan Research Group of Carabid Beetles 1989, Yahiro et al. 2001, Carabidological Society of Shiga 2003, Yahiro 2008), adult (Inoue 1953d), June (Inoue 1953d), larvae of Lymantria mathura, Zeuzera multistrigata, Camptoloma interioratum and Phalerodonta manleyi etc. (Inoue 1953d), larvae of Syntypistis punctatella (East Japan Research Group of Carabid Beetles 1989, Carabidological Society of Shiga 2003), larvae of lepidoptera and other insects (Suga 1981) hibernating after an active period in the autumn (Inoue 1953d), hibernating in soil (East Japan Research Group of Carabid Beetles 1989) |
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