| Ecology |
mountain (Kinki Research Group of Carabid Beetles 1979, Sota 1985c, Sota 1987b, Yahiro et al. 2001), hydric forest (Kinki Research Group of Carabid Beetles 1979), Quercus acutissima forest (Kinki Research Group of Carabid Beetles 1979), cedar forests (Togashi and Sugie 1994, Hiramatsu 2004), broad leaved deciduous forest (Togashi and Hashimoto 1994), edge of broad-leaved deciduous forest neighboring paddy fields (Togashi and Hashimoto 1994), beech forest (Hiramatsu 2004), grassland (Hiramatsu 2004), upland field (Hiramatsu 2004), adult (Sota 1985a, Sota 1985c) from spring to summer (Sota 1985a), from May to August (Sota 1985c), earthworm (Sota 1985a, Sota 1987b), earthworm (larva) (Sota 1985c) |
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