| Ecology |
mountain (Kinki Research Group of Carabid Beetles 1979, Sota 1987b, Yahiro et al. 2001, Yahiro 2008), from flatland to low mountain (Takakuwa et al. 2006), silvicolous (Ishitani 1996), pine forest (Kinki Research Group of Carabid Beetles 1979), Quercus acutissima forest (Kinki Research Group of Carabid Beetles 1979), bamboo forest (Kinki Research Group of Carabid Beetles 1979), wet forest floor (Kinki Research Group of Carabid Beetles 1979), Zelkova serrata forest (Kinki Research Group of Carabid Beetles 1979), Chamaecyparis obtusaforest (Kinki Research Group of Carabid Beetles 1979), maple forest (Kinki Research Group of Carabid Beetles 1979), beech forest (Kinki Research Group of Carabid Beetles 1979, Hiramatsu 2004), grassland (Hiramatsu 2004), orange field (Suga 1981), forest adjoinig agroecosystems (Yahiro et al. 1990), paddy field (Yahiro et al. 1992), broad leaved deciduous forest (Togashi and Hashimoto 1994), edge of broad-leaved deciduous forest neighboring paddy fields (Togashi and Hashimoto 1994), secondary forest of Alnus sieboldiana (Ishitani 1996), natural forests or near natural forests in mountain (Yahiro et al. 2001, Carabidological Society of Shiga 2003), dried secondary forest (Yahiro et al. 2001, Carabidological Society of Shiga 2003), cedar forests (Kinki Research Group of Carabid Beetles 1979, Togashi and Sugie 1994, Hiramatsu 2004), forest floor (Takakuwa et al. 2006), adult (Sota 1985a), from spring to summer (Sota 1985a), Megascolecoidea (Kinki Research Group of Carabid Beetles 1979), earthworm (Kinki Research Group of Carabid Beetles 1979, Sota 1985a, Sota 1987b), The larvae mainly eat Megascolecoidea (East Japan Research Group of Carabid Beetles 1989), collecting under lights (Kinki Research Group of Carabid Beetles 1979), hibernating in soil (Suga 1981), mainly hibernating in soil and rarely in decayed wood or under fallen tree (East Japan Research Group of Carabid Beetles 1989) |
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