| Ecology |
mountain (East Japan Research Group of Carabid Beetles 1989, Kimoto and Yasuda 1995), herbosa (East Japan Research Group of Carabid Beetles 1989), seaboard herbosa (East Japan Research Group of Carabid Beetles 1989, Kimoto and Yasuda 1995), broad-leaved deciduous forest (East Japan Research Group of Carabid Beetles 1989), mixed forest (East Japan Research Group of Carabid Beetles 1989), forest edge (East Japan Research Group of Carabid Beetles 1989), forest zone (Kimoto and Yasuda 1995, Hori 2001), Pinus pumila community (Kimoto and Yasuda 1995), snowy wet grassland of the alpine region (Kimoto and Yasuda 1990, Kimoto and Yasuda 1995), Betula ermanii forest (Kimoto and Yasuda 1995), coniferous forest (Kimoto and Yasuda 1995), higher mixed forest (Kimoto and Yasuda 1990, Kimoto and Yasuda 1995), lower mixed forest (Kimoto and Yasuda 1995), altherbosa (Kimoto and Yasuda 1995), Abies sachalinensis forest (Kimoto and Yasuda 1995, Hori 2003), Sasa kurilensis community (Kimoto and Yasuda 1995), Polygonum sachalinense community (Kimoto and Yasuda 1995), seaboard herbosa (Kimoto and Yasuda 1995), Quercus crispula forest (Kimoto and Yasuda 1995), generally larva, and adult (Sota 1985a), from late April to early May in lowland (East Japan Research Group of Carabid Beetles 1989), June in upland and cold region (East Japan Research Group of Carabid Beetles 1989), snail (Sota 1985a, East Japan Research Group of Carabid Beetles 1989), seeds of Morus australis (East Japan Research Group of Carabid Beetles 1989), Parapodisma mikado (East Japan Research Group of Carabid Beetles 1989) |
| Remarks |
subspecies: ssp. gehinii, ssp. aereicollis, ssp. konsenensis, ssp. radiatocostatus, ssp. sapporensis, ssp. shimizui, ssp. manoianus, ssp. munakatai, ssp. nishijimai |
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