| Ecology |
forest (Inoue 1952s), plantation of Larix leptolepis saplings (Higashi et al. 1983), moor land (East Japan Research Group of Carabid Beetles 1989), wilderness (East Japan Research Group of Carabid Beetles 1989), broad-leaved secondary forest (East Japan Research Group of Carabid Beetles 1989), coniferous forest (East Japan Research Group of Carabid Beetles 1989, Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), primary forest (East Japan Research Group of Carabid Beetles 1989), snowy wet grassland of the alpine region (Kimoto and Yasuda 1990, Kimoto and Yasuda 1994, Kimoto and Yasuda 1995), Betula ermanii forest (Kimoto and Yasuda 1990, Kimoto and Yasuda 1991, Kimoto and Yasuda 1995), Picea jezoensis and Betula ermanii forest (Kimoto and Yasuda 1991), higher mixed forest (Kimoto and Yasuda 1994), lower mixed forest (Kimoto and Yasuda 1995), Pinus pumila community (Kimoto and Yasuda 1995), altherbosa (Kimoto and Yasuda 1995), Abies sachalinensis forest (Kimoto and Yasuda 1995), Sasa kurilensis community (Kimoto and Yasuda 1995), Polygonum sachalinense community (Kimoto and Yasuda 1995), mixed forest (Hori 2003), park forest (Hori 2003), riverside forest (Hori 2003), adult and larva (Inoue 1952s, East Japan Research Group of Carabid Beetles 1989), June (Inoue 1952s), carnivorous (Inoue 1952s), snail (Inoue 1952s), Lepidopteran larva (Inoue 1952s), the adults and larvae are snail feeder (East Japan Research Group of Carabid Beetles 1989), Discus pauper (East Japan Research Group of Carabid Beetles 1989), adults form hibernating assemblage (Inoue 1952s), aestivation？ (Inoue 1952s), hibernating in decayed fallen tree (East Japan Research Group of Carabid Beetles 1989) |
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