| Ecology |
mountain (Sota 1987b, Sota 1985c, Yahiro et al. 2001, Carabidological Society of Shiga 2003, Yahiro 2008), pasania forest (Kinki Research Group of Carabid Beetles 1979), beech forest (Kinki Research Group of Carabid Beetles 1979), Chamaecyparis obtusa forest (Kinki Research Group of Carabid Beetles 1979), ridge of dried low mountain (East Japan Research Group of Carabid Beetles 1989), beech area and coniferous forest zone in high altitude (East Japan Research Group of Carabid Beetles 1989), Cryptomeria japonica forest (Kinki Research Group of Carabid Beetles 1979, Togashi and Sugie 1994), natural forests or near natural forests in mountain and dried secondary forest (Kinki Research Group of Carabid Beetles 1979, Yahiro et al. 2001, Carabidological Society of Shiga 2003), adult (Sota 1985a, Sota 1985c), from spring to summer (Sota 1985a), from May to August (Sota 1985c), earthworm (larva) (Sota 1985a, Sota 1985c, Sota 1987b, East Japan Research Group of Carabid Beetles 1989) |
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