Ostracod Research at the Lake Biwa Museum, Japan

Robin James Smith

Japan

Total = 111 species.

= species currently only known from Japan (47 species).
= species invasive to Japan (seven species).

Order Podocopida Sars, 1866
Suborder Cypridocopina Jones, 1901
Superfamily Cypridoidea Baird, 1845
Family Candonidae Kaufmann, 1900
Subfamily Candoninae Kaufmann, 1900
Genus Candona Baird, 1845
Candona candida (O. F. Müller, 1776)
Candona subgibba Sars, 1926

Genus Candonopsis Vávra, 1891
Candonopsis kingsleii (Brady & Robertson, 1870)
Candonopsis tenuis (Brady, 1886)

Genus Cryptocandona Kaufmann, 1900
Cryptocandona brehmi (Klie, 1934)
Cryptocandona tsukagoshii Smith, 2011

Genus Fabaeformiscandona Krstić, 1972
Fabaeformiscandona akaina Smith & Janz, 2008
Fabaeformiscandona condylea Smith & Janz, 2008
Fabaeformiscandona dolabella Smith & Janz, 2008
Fabaeformiscandona kushiroensis Hiruta & Hiruta, 2015
Fabaeformiscandona myllaina Smith & Kamiya, 2007
Fabaeformiscandona nishinoae Smith & Janz, 2008
Fabaeformiscandona okuboi Smith & Janz, 2008
Fabaeformiscandona paterea Smith & Janz, 2008
Fabaeformiscandona pedana Smith & Janz, 2008
Fabaeformiscandona subacuta (Yang, 1982)
Fabaeformiscandona tora Smith & Kamiya, 2007
Fabaeformiscandona velifera Smith & Janz, 2008
Fabaeformiscandona yajimae Smith & Janz, 2008

Genus Pseudocandona Kaufmann, 1900
Pseudocandona abei Smith & Janz, 2008
Pseudocandona atmeta Smith & Kamiya, 2015
Pseudocandona becca Smith & Kamiya, 2015
Pseudocandona falcula Smith & Kamiya, 2015
Pseudocandona pratensis (Hartwig, 1901)
Pseudocandona tenuirostris Hiruta & Mawatari, 2013

Genus Schellencandona Meisch, 1996
Schellencandona yakushimaensis Smith & Kamiya, 2006

Genus Undulacandona Smith, 2011
Undulacandona colymba Smith & Kamiya, 2015
Undulacandona spinula Smith, 2011

Subfamily Cyclocypridinae Kaufmann, 1900
Genus Cyclocypris Brady & Norman, 1889
Cyclocypris diebeli Absolon, 1973
Cyclocypris ovum (Jurine, 1820)

Genus Cypria Zenker, 1854
Cypria exsculpta (Fischer, 1855)
Cypria konishii Smith & Kamiya, 2006
Cypria matzkeae Smith & Janz, 2008

Genus Physocypria Vávra, 1897
Physocypria biwaensis (Okubo, 1990)
Physocypria nipponica Okubo, 1990

Subfamily Paracypridinae Sars, 1923
Genus Dolerocypria Tressler, 1937
Dolerocypria mukaishimensis Okubo, 1980

Genus Paracypria Sars, 1910
Paracypria adnata Smith & Kamiya, 2006
Paracypria inujimensis (Okubo, 1980)

Family Cyprididae Baird, 1845
Subfamily Cyprettinae Hartmann, 1971
Genus Cypretta Vávra, 1895
Cypretta seurati Gauthier, 1929

Subfamily Cypricercinae McKenzie, 1971
Genus Bradleycypris McKenzie, 1982
Bradleycypris vittata (Sars, 1903)

Genus Bradleytriebella Savatenalinton & Martens, 2009
Bradleytriebella lineata (Victor & Fernando, 1981)
Bradleytriebella tuberculata (Hartmann, 1964)

Genus Pseudostrandesia Savatenalinton and Martens, 2009
Pseudostrandesia tenebrarum Smith & Ozawa, 2021 ?

Genus Tanycypris Triebel, 1959
Tanycypris alfonsi Nagler, Geist & Matzke-Karasz, 2014

Subfamily Cypridinae Baird, 1845
Genus Chlamydotheca Saussure, 1858
Chlamydotheca arcuata (Sars, 1901)

Genus Cypris O.F. Müller, 1776
Cypris granulata Daday, 1898
Cypris maculosa Bronstein, 1925

Subfamily Cypridopsinae Kaufmann, 1900
Genus Cavernocypris Hartmann, 1964
Cavernocypris cavernosa Smith, 2011
Cavernocypris hokkaiensis Munakata et al., 2022

Genus Cypridopsis Brady, 1867
Cypridopsis nigrovittata Okubo, 1990
Cypridopsis uenoi Brehm, 1933
Cypridopsis vidua (O. F. Müller, 1776)

Genus Potamocypris Brady, 1870
Potamocypris kusuokai Smith, 2023
Potamocypris mastigophora (Methuen, 1910)

Genus Sarscypridopsis McKenzie, 1977
Sarscypridopsis denticulata Smith & Ozawa, 2023 ?

Genus Siamopsis Savatenalinton, 2017
Siamopsis sudzukii (Okubo, 1992)

Subfamily Cyprinotinae Bronstein, 1947
Genus Cyprinotus Brady, 1886
Cyprinotus cassidula Smith & Chang, 2020
Cyprinotus kimberleyensis McKenzie, 1966

Genus Hemicypris Sars, 1903
Hemicypris barbadensis Broodbakker, 1983
Hemicypris kaufmanni (Vávra, 1906)
Hemicypris kibiensis Okubo, 1990
Hemicypris mizunoi Okubo, 1990
Hemicypris okayamenesis Okubo, 2004 nomen nudum
Hemicypris posterotruncata Bate, 1970 ?

Genus Heterocypris Claus, 1892
Heterocypris auricularis Zhai & Zhao, 2014
Heterocypris barbara (Gauthier & Brehm, 1928)
Heterocypris incongruens (Ramdohr, 1808)
Heterocypris savatenalintonae Smith & Chang, 2020
Heterocypris spadix Munakata et al., 2021

Subfamily Dolerocypridinae Triebel, 1961
Genus Dolerocypris Kaufmann, 1900
Dolerocypris fasciata (O. F. Müller, 1776)
Dolerocypris ikeyai Smith & Kamiya, 2006
Dolerocypris sinensis Sars, 1903

Subfamily Eucypridinae Bronstein, 1947
Genus Eucypris Vávra, 1891
Eucypris pigra (Fischer, 1851)

Subfamily Herpetocypridinae Kaufmann, 1900
Genus Candonocypris Sars, 1896
Candonocypris novaezelandiae (Baird, 1843)
Candonocypris caledonica (Méhes, 1939)

Genus Chrissia Hartmann, 1957
Chrissia formosa (Klie, 1938)
Chrissia vittata Okubo, 1974

Genus Herpetocypris Brady & Norman, 1889
Herpetocypris chevreuxi (Sars, 1896)

Genus Ilyodromus Sars, 1894
Ilyodromus intermedius Okubo, 2011

Genus Stenocypris Sars, 1889
Stenocypris hirutai Smith & Kamiya, 2006
Stenocypris hislopi Ferguson, 1969
Stenocypris major (Baird, 1859)
Stenocypris malayica Victor & Fernando, 1981
Stenocypris tsukagoshii Smith & Kamiya, 2006
Stenocypris viridis Okubo, 1990

Subfamily Scottinae Bronstein, 1947
Genus Scottia Brady & Norman, 1889
Scottia birigida Smith et al., 2002
Scottia pseudobrowniana Kempf, 1971

Family Ilyocyprididae Kaufmann, 1900
Subfamily Ilyocypridinae Kaufmann, 1900
Genus Ilyocypris Brady & Norman, 1889
Ilyocypris angulata Sars, 1903
Ilyocypris dentifera Sars, 1903
Ilyocypris incus Smith et al., 2019
Ilyocypris japonica Okubo, 1990
Ilyocypris salebrosa Stepanaitys, 1960

Family Notodromadidae Kaufmann, 1900
Subfamily Notodromadinae Kaufmann, 1900
Genus Notodromas Lilljeborg, 1853
Notodromas trulla Smith & Kamiya, 2014

Suborder Cytherocopina
Superfamily Cytheroidea Baird, 1850
Family Cytherideidae Sars, 1925
Genus Cytherissa Sars, 1925
Cytherissa lacustris (Sars, 1863)

Family Cytheromatidae Elofson, 1939
Genus Microloxoconcha Hartmann, 1954
Microloxoconcha semicircularis Smith & Chang, 2022

Family Limnocytheridae Klie, 1938
Subfamily Limnocytherinae Klie, 1938
Genus Limnocythere Brady, 1868
Limnocythere cyphoma Smith & Janz, 2009
Limnocythere fude Smith & Janz, 2009
Limnocythere kamiyai Smith & Janz, 2009
Limnocythere levigatus Smith & Janz, 2009
Limnocythere stationis Várva, 1891

Subfamily Timiriaseviinae Mandelstam, 1960
Genus Metacypris Brady & Robertson, 1870
Metacypris digitiformis Smith & Hiruta, 2004

Family Entocytheridae Hoff, 1942
Subfamily Entocytherinae Hoff, 1942
Genus Ankylocythere Hart, 1962
Ankylocythere sinuosa (Rioja, 1942)

Genus Uncinocythere Hart, 1962
Uncinocythere occidentalis (Kozloff & Whitman, 1954)

Suborder Darwinulocopina Brady & Robertson, 1885
Superfamily Darwinuloidea Brady & Robertson, 1885
Family Darwinulidae Brady & Robertson, 1885
Genus Darwinula Brady & Robertson, 1885
Darwinula stevensoni (Brady & Robertson, 1870)

Genus Microdarwinula Danielopol, 1968
Microdarwinula zimmeri (Menzel, 1916)

Genus Penthesilenula Rossetti & Martens, 1998
Penthesilenula brasiliensis (Pinto & Kotzian, 1961)

Genus Vestalenula Rossetti & Martens, 1998
Vestalenula cornelia Smith et al., 2006
Vestalenula cylindrica (Straub, 1952)
Vestalenula gravata Smith & Chang, 2022
Vestalenula lundi (Neale & Victor, 1978)
Vestalenula molopoensis (Martens & Rossetti, 1997)

Korea

Total = 59 species.

= species currently only known from Korea (20 species).

Order Podocopida Sars, 1866
Suborder Cypridocopina Jones, 1901
Superfamily Cypridoidea Baird, 1845
Family Candonidae Kaufmann, 1900
Subfamily Candoninae Kaufmann, 1900
Genus Candona Baird, 1845
Candona quasiakaina Karanovic & Lee, 2012
Candona sillae Karanovic & Lee, 2012

Genus Candonopsis Vávra, 1891
Candonopsis transgrediens Brehm, 1923

Genus Cryptocandona Kaufmann, 1900
Cryptocandona brehmi (Klie, 1934)
Cryptocandona smithi Karanovic & Lee, 2012
Cryptocandona tsukagoshii Smith, 2011

Genus Earicandona Karanovic, 2013
Earicandona mounchyon Karanovic, 2013

Genus Fabaeformiscandona Krstić, 1972
Fabaeformiscandona koreana Karanovic & Lee, 2012
Fabaeformiscandona subacuta (Yang, 1982)

Genus Nannocandona Ekman, 1914
Nannocandona schornikovi Karanovic, Sidorov & Marmonier, 2015

Genus Pseudocandona Kaufmann, 1900
Pseudocandona choi (Karanovic & Lee, 2012)
Pseudocandona morimotoi (McKenzie, 1972)
Pseudocandona pratensis (Hartwig, 1901) (as Typhlocypris pratensis (Hartwig, 1901))

Genus Schellencandona Meisch, 1996
Schellencandona tea Karanovic & Lee, 2012

Genus Undulacandona Smith, 2011
Undulacandona aeolus Karanovic & Cho, 2017
Undulacandona andromeda Karanovic & Cho, 2017
Undulacandona cetus Karanovic & Cho, 2017
Undulacandona koreana Karanovic & Cho, 2017

Subfamily Cyclocypridinae Kaufmann, 1900
Genus Cyclocypris Brady & Norman, 1889
Cyclocypris anacola Smith, Lee & Chang, 2014

Genus Cypria Zenker, 1854
Cypria asiatica Karanovic & Lee, 2012

Genus Physocypria Vávra, 1897
Physocypria kraepelini Müller, 1903
Physocypria nipponica Okubo, 1990

Subfamily Paracypridinae Sars, 1923
Genus Dolerocypria Tressler, 1937
Dolerocypria mukaishimensis Okubo, 1980

Family Cyprididae Baird, 1845
Subfamily Cyprettinae Hartmann, 1971
Genus Cypretta Vávra, 1895
Cypretta karanovicae Smith et al., 2014
Cypretta seurati Gauthier, 1929

Subfamily Cypricercinae McKenzie, 1971
Genus Bradleycypris McKenzie, 1982
Bradleycypris vittata (Sars, 1903)

Genus Bradleytriebella Savatenalinton & Martens, 2009
Bradleytriebella tuberculata (Hartmann, 1964)

Genus Strandesia Stuhlmann, 1888
Strandesia flavescens Klie, 1932

Genus Tanycypris Triebel, 1959
Tanycypris alfonsi Nagler, Geist & Matzke-Karasz, 2014
Tanycypris centa Chang, Lee & Smith, 2012

Subfamily Cypridopsinae Kaufmann, 1900
Genus Cavernocypris Hartmann, 1964
Cavernocypris coreana coreana (McKenzie, 1972)
Cavernocypris coreana elongata (McKenzie, 1972)
Cavernocypris danielopoli Smith & Kamiya, 2017

Genus Cypridopsis Brady, 1867
Cypridopsis vidua (O. F. Müller, 1776)

Genus Potamocypris Brady, 1870
Potamocypris mastigophora (Methuen, 1910)
Potamocypris variegata (Brady & Norman, 1889)

Subfamily Cyprinotinae Bronstein, 1947
Genus Cyprinotus Brady, 1886
Cyprinotus cassidula Smith & Chang 2020

Genus Hemicypris Sars, 1903
Hemicypris posterotruncata Bate, 1970

Genus Heterocypris Claus, 1892
Heterocypris incongruens (Ramdohr, 1808)
Heterocypris salina (Brady, 1868)

Subfamily Dolerocypridinae Triebel, 1961
Genus Dolerocypris Kaufmann, 1900
Dolerocypris fasciata (O. F. Müller, 1776)
Dolerocypris ikeyai Smith & Kamiya, 2006
Dolerocypris sinensis Sars, 1903

Subfamily Eucypridinae Bronstein, 1947
Genus Eucypris Vávra, 1891
Eucypris pigra (Fischer, 1851)

Subfamily Herpetocypridinae Kaufmann, 1900
Genus Stenocypris Sars, 1889
Stenocypris hirutai Smith & Kamiya, 2006
Stenocypris hislopi Ferguson, 1969

Subfamily Scottinae Bronstein, 1947
Genus Scottia Brady & Norman, 1889
Scottia birigida Smith et al, 2002
Scottia pseudobrowniana Kempf, 1971

Family Ilyocyprididae Kaufmann, 1900
Subfamily Ilyocypridinae Kaufmann, 1900
Genus Ilyocypris Brady & Norman, 1889
Ilyocypris bradyi Sars, 1890
Ilyocypris dentifera Sars, 1903
Ilyocypris hanguk Karanovic & Lee, 2013
Ilyocypris incus Smith et al., 2019
Ilyocypris japonica Okubo, 1990
Ilyocypris salebrosa Stepanaitys, 1960
Ilyocypris subpulchra Yang, 1982

Suborder Cytherocopina
Superfamily Cytheroidea Baird, 1850
Family Limnocytheridae Klie, 1938
Subfamily Limnocytherinae Klie, 1938
Genus Limnocythere Brady, 1868
Limnocythere stationis Vavra, 1891

Subfamily Timiriaseviinae Mandelstam, 1960
Genus Frambocythere Colin, 1980
Frambocythere relicta Smith et al., 2012

Superfamily Terrestricytheroidea Schornikov, 1969
Family Terrestricytheridae Schornikov, 1969
Genus Terrestricythere Schornikov, 1969

Terrestricythere ivanovae Schornikov, 1969

Suborder Darwinulocopina Brady & Robertson, 1885
Superfamily Darwinuloidea Brady & Robertson, 1885
Family Darwinulidae Brady & Robertson, 1885
Genus Vestalenula Rossetti & Martens, 1998
Vestalenula cylindrica (Straub, 1952)

Listed are those that have been identified down to the species level (i.e. excludes Candona sp. 1, Cypridopsis cf. vidua etc).